B-17 with exotic painting

Your link is broken it should be this...

http://aviacaoemfloripa.blogspot.com.br/2011/01/boeing-b-17-flying-fortress-spotted-cow.html

The answer to your questions are:
1) Yes it entered service and flew 61 missions with the 384th Bomb Group.
2) The aircraft as you see it there was painted that way as an "assembly aircraft" it took off and positioned itself for other aircraft in the bomb group to "assemble" around.
3) Each assembly aircraft was painted in a different pattern so that it could be easily spotted by those forming up around it.
 
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